The film begins with a narration by Io, explaining that there were originally three Gods: Zeus, Poseidon, and Hades. The Gods defeated their greatest enemies, the Titans, with the help of Hades's creation, the Kraken. After the Titans' defeat, Zeus created humans and ruled them while Poseidon ruled the sea, but Hades, having been deceived by his brother Zeus was forced to rule the Underworld, constantly plotting to take Zeus's throne for himself.
A fisherman, Spyros, finds a coffin adrift in the sea, discovering a baby, Perseus, and his dead mother, Danaë, inside. Spyros decides to raise Perseus as his own. Years later, Perseus and his family are fishing when they witness a group of soldiers from Argos destroying a statue of Zeus and declaring war against the Gods. Hades appears and commands harpies to massacre the soldiers. The harpies then destroy Perseus's family's fishing boat. Perseus tries to save his family but to no avail, the surviving soldiers take Perseus back to Argos.
During a feast for the returning soldiers King Cepheus and Queen Cassiopeia of Argos compare themselves and their daughter, Andromeda, to the Gods. Andromeda is against fighting the Gods, and is protesting when Hades appears and kills the remaining soldiers. Perseus is unaffected however, and Hades realizes Perseus is a demigod and reveals Zeus is Perseus's father. Hades kills the queen, and threatens that if Princess Andromeda is not sacrificed to the Kraken, Argos will be destroyed in 10 days.
The king seeks the help of Perseus. Perseus initially refuses, but decides to help when he meets Io, a woman who does not age as punishment for refusing to be seduced by a God. Io has watched over Perseus from the time he was born, and assures him he can have his revenge against Hades if he helps Argos in its time of need. Perseus and Io embark on a quest to find the Stygian Witches. Hades enlists Acrisius, the former king of Argos who was married to Danaë when Zeus impregnated her with Perseus. Acrisius was leading men to attack Olympus, and while other gods wanted to annihilate Acrisius, Zeus wanted to make an example of him. He snuck into Danaë's bed in the form of Acrisius and impregnated her. Acrisius declared that the baby could not be born, and set Danaë and the baby Perseus adrift in their coffin; for this act, Zeus struck Acrisius with lightning, leaving him hideously disfigured. Hades imbues him with powers to aid him in killing Perseus, and Acrisius takes the name of Calibos. While in the woods, Perseus and his men discover a sword forged in Olympus that will only display power if it is wielded by Perseus. Perseus also discovers a winged horse called Pegasus. Perseus refuses to use either of these gifts, as he does not wish to be a God. Calibos attacks the group led by Perseus. They severe the hand of Calibos, but Calibos' blood forms giant scorpions from the sand, and Perseus and his group are attacked. Though they slay several scorpions, most of the group are slain, and the survivors surrounded by more of the monsters. They are saved by the Djinn, sand-demons with magical powers who tame the scorpions. Perseus and the Djinn agree to cooperate, since the Djinn wish to see the Gods defeated.
At the Garden of Stygia, the Stygian Witches tell Perseus that the head of Medusa could kill the Kraken, but that Perseus will die in the process. After leaving the witches, Perseus is visited by Zeus who offers him asylum on Mount Olympus. He refuses. Zeus gives him instead a golden drachma. Perseus uses the coin to bribe Charon for passage into the Underworld and enters Medusa's lair. Although four of Perseus' remaining soldiers fight hard to stay alive, one by one they are turned to stone by her gaze. Draco, one of the soldiers with Perseus, allows himself to be turned to stone to give Perseus a clear shot at Medusa's head. Perseus kills Medusa, but Calibos returns and kills Io. Perseus engages Calibos in combat and kills him, using the sword from Olympus to stab him in the heart and turn him back into Acrisius in human form. With his final breaths, Acrisius tells Perseus to never become a God. Perseus rides Pegasus back to Argos with Medusa's head to find the citizens of Argos planning to sacrifice Andromeda to the Kraken, who has been summoned by Hades. Hades reveals to Zeus the destruction of Argos will give him enough power to overthrow the other Olympians. Zeus attempts to banish Hades back to the Underworld, but Hades overpowers him. Zeus then reveals to Hades that Perseus is still alive and is in Argos. Hades leaves to oversee the Kraken's attack and ensure his victory.
Perseus returns to Argos, but Hades sends harpies to kill him. Perseus defeats the creatures sent by Hades and uses the head of Medusa to turn the Kraken into stone. Hades arrives just in time to see his creation die. Perseus draws his sword, but Hades reminds him that he is a God, and therefore cannot be killed. Perseus replies that though Hades can live forever, it will not be in the world of men and throws his sword at Hades, charged with a bolt of lightning from Zeus that banishes Hades to the Underworld. After saving Argos from destruction, Andromeda suggests that Perseus become king and rule Argos at her side, but he declines.
Zeus appears before Perseus again and offers to make him a God, but he refuses. Zeus warns Perseus that Hades will return to rule the world in darkness when mankind is fearful. Since Perseus is intent to stay on Earth, Zeus resurrects Io, and the two embrace while Pegasus flies above them.
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